Development of non-insulin-dependent diabetes mellitus in the double knockout mice with disruption of insulin receptor substrate-1 and β cell glucokinase genes: Genetic reconstitution of diabetes as a polygenic disease

Yasuo Terauchi, Keiji Iwamoto, Hiroyuki Tamemoto, Kajuro Komeda, Chikara Ishii, Yasunori Kanazawa, Nahoko Asanuma, Toru Aizawa, Yasuo Akanuma, Kazuki Yasuda, Tatsuhiko Kodama, Kazuyuki Tobe, Yoshio Yazaki, Takashi Kadowaki*

*Corresponding author for this work

Research output: Contribution to journalArticlepeer-review

132 Scopus citations

Abstract

Non-insulin-dependent diabetes mellitus (NIDDM) is considered a polygenic disorder in which insulin resistance and insulin secretory defect are the major etiologic factors. Homozygous mice with insulin receptor substrate-1 (IRS-1) gene knockout showed normal glucose tolerance associated with insulin resistance and compensatory hyperinsulinemia. Heterozygous mice with β cell glucokinase (GK) gene knockout showed impaired glucose tolerance due to decreased insulin secretion to glucose. To elucidate the interplay between insulin resistance and insulin secretory defect for the development of NIDDM, we generated double knockout mice with disruption of IRS-1 and β cell GK genes by crossing the mice with each of the single gene knockout. The double knockout mice developed overt diabetes. Blood glucose levels 120 min after intraperitoneal glucose load (1.5 mg/g body wt) were 108±24 (wild type), 95±26 (IRS-1 knockout), 159±68 (GK knockout), and 210±38 (double knockout) mg/dl (mean±SD) (double versus wild type, IRS-1, or GK; P < 0.01). The double knockout mice showed fasting hyperinsulinemia and selective hyperplasia of the β cells as the IRS-1 knockout mice (fasting insulin levels: 0.38±0.30 [double knockout], 0.35±0.27 [IRS-1 knockout] versus 0.25±0.12 [wild type] ng/ml) (proportion of areas of insulin-positive cells to the pancreas: 1.18±0.68%; P < 0.01 [double knockout], 1.20±0.93%; P < 0.05 [IRS-1 knockout] versus 0.54±0.26% [wild type]), but impaired insulin secretion to glucose (the ratio of increment of insulin to that of glucose during the first 30 min after load: 31 [double knockout] versus 163 [wild type] or 183 [IRS-1 knockout] ng insulin/mg glucose x 103). In conclusion, the genetic abnormalities, each of which is nondiabetogenic by itself, cause overt diabetes if they coexist. This report provides the first genetic reconstitution of NIDDM as a polygenic disorder in mice.

Original languageEnglish
Pages (from-to)861-866
Number of pages6
JournalJournal of Clinical Investigation
Volume99
Issue number5
DOIs
StatePublished - 1997/03/01

Keywords

  • glucokinase
  • insulin receptor substrate-1
  • insulin resistance
  • insulin secretory dysfunction
  • non-insulin-dependent diabetes mellitus

ASJC Scopus subject areas

  • General Medicine

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